Updated for 2004 Graphs only.
The following has been updated to include in the charts and graphs the
data for 2001. Only the charts have been updated to 2004 except for the
Calanoids, they've been graphed but looking at how the three months data
fits I don't think the graphs add any clarity so I've decided not to do
them for the the Cyclopoids and Nauplii.
MONTHLY AVERAGES FOR COPEPODS
At times I think it is totally ludicrous to think the samples taken have
any significance at all. I am taking a paltry bucket of water from the
Lake and expecting a little 12.5 liter bit of water to represent what is
occurring in the Lake's 100 billion liter biologic community. The only
reason I can even have the temerity to suggest such a thing is that all
the other researcher have done the same. Granted they may have taken a
couple hundred more liters in their samples, but this is the merest pittance
in difference when compare with the volume of water flowing through the
Lake. Besides where my samples lack individual size the gathering of samples
has been continually carried on longer than any sampling I know of
in the Lake. So while I don't hold the following numbers in high esteem
I assign even less worth to most studies done in the past which may have
only covered one year or had only 20 or 30 samples per year.
So let us crunch some
numbers. Remember I'm an amateur, with but a touch of knowledge of biology
and even less knowledge of statistics. The numbers generated below were
taken from 12.5 liters samples. The samples were taken twice a week since
1992 (though I have not been religious about getting the samples, there
are a few weeks were only one sample was taken and other weeks over the
years when no sample was taken at all). When I first started this project
I felt compelled to attempt to get samples year round under any conditions,
this became a mission from God. After a couple of years of pounding through
the ice to get to the water I gave up on taking winter samples so the continuous
range of samples really extends only from April to December.
Addressing
the question of methodology; I've had to make up my own method of counting
the creatures. First I walk out on the stone jetties at the Veterans Memorial
Park, tie a rope to a plastic bucket that I've put a rock in, toss it out
and let it sink. Twelve point five liters of water from the buckets is
filter through two coffee filters until about 50 to 100 ml. are left. This
water is than pour into a small dish at about 10 to 20 ml. at a time and
the animals counted using a dissecting microscope. I only count animals
greater than. 05 mm. In case anyone is wondering why 12.5 liters is the
amount of water used in the sample let me explain. I was going to use ten
liters as the sample size so I figured I would measure out ten liters using
two liter Pepsi bottles, trouble is a two liter bottle holds more than
two liters when filled to the top, they hold about .5 liters more. I didn't
realize this for several years into the study when I measured the water
again using a measuring cup marked in liters. Rather than changing the
sample size I just continued to take 12.5-liter samples.
COPEPODS
FROM 1993 TO 2004, AVON POINT
To begin: below is a graph giving the overall all view of what the samples
showed over the seven years. That graph is arrived at by taking the average
number of animals found in the samples monthly and than taking an average
of this. THE GRAPH ONLY COVERS THE MONTHS FROM APRIL TO DECEMBER.
The graph glosses over in its
simplicity a whole range of problems in trying to interpret the data as
can be seen by studying the charts presented below.
Below are a lot of figures, which
may look impressive but I caution not to read into them more accuracy than
is there. I count living animals, which is hard to do since they swim fast.
The more animals there were in a sample the less accurate was the count.
It would have been better if I had broken out the different species of
copepods when I counted them, but I only have so much time to devote to
this aspect of the Lake so this is the best I could manage.
CALANOID COPEPODS
Monthly Averages for Calanoid Copepods
The following data is based on 12.5-liter samples
taken twice a week. All the samples were taken in the morning from the
same location.
Calanoid Copepods 1993 to 2004
1993 1994
1995 1996
1997 1998 1999
2000
2001 2003
2004
Jan
17.2
Feb
.3
Mar.
2.2 2.6
.9 1.6
.25
Apr.
7.7 3.7
3.5 4.5
24.1 2.9
18.4
3.6 7.3
7
May
104.3 84.8
37 40.5
11.6 15.2
19.7
11.2 21
32.7
June 12.8
71.4 56.7
60.7 *144 27.8
26.7 15.2
6.7 28.1
64
July 4.9
22.4 2.4
28.4 8.5
4.6
8.1 13.2
8.4 47.2
59
Aug 4
16.7 5
28.1 7.1
2.6 6.1
6.9
16.1
Sept. 26.6
32.1 13.6
6.8 11
12
11.9 32.7
1.9
Oct 15.6
11 9.3
34.6 6.3
21
3.8
8.3 25.9
Nov 9.1
9.9 5.3
2.9 4.9
14.3 12.4
2.8 10.5
Dec 5.6
6.3 7.2
5.6 .8
20.1 5.5
35.7
Average of the Calanoids Apr. thru
Dec. 1994 1995 1996
1997 1998 1999 2000
2001
31.3 20.9 23.04 25.3
15.34 10.29 17
10.5
*Except for the extraordinary
number of animals found on one sampling date the number of calanoid copepods
would have shown a marked decline in 1997. That date was June 15. The sample
had the extraordinary number of 880 calanoids in the 12.5 liter sample.
The total number of calanoids found for the entire year in about 50 samples
minus the June 15 date was 1130, thus the one sample had more animals than
all the samples for the entire year. I have belabored the point of the
non randomness of the zooplankton in the 1996 report and so will not cover
that ground again. What I want to point out here is that if the June 15
sample is dropped from the reporting than the copepods had a steep drop
in their numbers in 1997 and the average for the month of June would be
52 per sample rather than 144 per sample. The average for the year would
drop from 25.3 per sample to 11.4 per sample. Whether this dropping of
the sample from the total counts is valid I leave up to the reader but
if the June 15 sample is ignored than the calanoids have had a major decline
in 1997.
There is also a statistical anomaly in the December 2000 sample of 12/10.
This sample had 99 calanoids in it which skewed upward the whole average
greatly. Below are the numbers for the whole month. A note about the few
December samples taken: this has been the first year for quite a while
that the near shore area of the Lake has frozen over this early, which
is why so few sample were taken.
12/3 13.00
12/6 20.00
12/10 99.00
12/13 7.00
Not only were the Calanoids
found in high numbers but so were (to a lesser but significant extent)
the Cyclopoids, as will be seen below. Again, oh what to do with these
anomalies?
CALANOID COPEPODS 1993 TO 1998
CYCLOPOID COPEPODS
The table does not include the cyclopoid T. Mexicanus
pranis. This small copepod was found very sporadically in the samples until
this year. Though when it was found it could be found in large numbers.
Average for the years 1994 1995
1996 1997 1998 1999
2000 2001
Apr. to Dec.
18.3 34.2 24.5
10.3 12.8 29.7
13.6 62.2
There were a total of 29 plankton water samples taken in 2004.
The samples were taken between April 2 and
July 12. There were in April 8 samples taken
May 8 "
" The July samples present
a problem in that the three samples
June 9 "
" taken at the beginning
of the month had a high animal count
July 4 "
" which if averaged
in to what would normally occur in July as
the number of animals tapered off would give a much lower
average number of animals.
Below is just the current data for more details on the methods
and meanings of the numbers here listed see the links below.
Comparison of Cyclopoids Copepods from 1993 to 2004
Monthly Averages 12.5 liter samples
1993 1994
1995 1996 1997
1998 1999 2000
2001 2003
2004
April
8.6 9.8
3 .3
2.1 2.4
1.9 5.4
4.5 2
May
38.3 56.8 50.3
8.9 34.7 24.6
24.5 228.7
9.1 10.9
June
51.5 38.8 182
109.4 35.4 59.1
89.2 45.2
205.7
253.4 114.3
July
25 26.6
19.6 17.1 30.2
6.3 111.4* 16.8
1.8 69.4
86.8
Aug.
38.5 13.7 19.1
5 7.7
3.7 9.4
4.6 7.7
Sept.
17.3 12.5
7.1 17.3 2.9
1.2 21.9
10.1 5.4
Oct.
8.6 13.1
3.8 8.9
2.3 .4
3.2 1.4
39.9
Nov.
12.9 7.7
5.7 1.5
4.3 1.6
1.3 3.5
2.7
Dec.
10.7 5.1
4.4 7.7
1.1 3.3
4.2 14.3
*The average for July,
1999 is skewed by one sampling date (July 14) with a total of 638 cyclopoids
in the sample. Eliminating the one sample from the average for the month
gives a new monthly average of 23.7.A further note on this sample. After
taking the plankton sample from off the jetty I went out in a boat about
a 1/2 mile from shore to get the benthic sample in about 15 feet of water.
The number of cyclopoids in that 5-liter sample was 184. Multiplying this
by 2.5 to make it equal the plankton sample gives a total of 460. Meaning
this massing of cyclopoids in the area probably covered a large area. The
cyclopoids totals taken from each sample for the month of July were:
July 5
45
8 21
11 1
14 638
21 60
25 7
28 8
Avg 111.4
The December of 2000 samples, as mention above, had a day of aberration
in the samples. Below are all the samples for the month.
12/3 6.00
12/6 15.00
12/10 32.00
12/13 6.00
Below is a graph of the cyclopoids found
in the 12.5 liter samples again in for the months of May and June from
1993 to 2000.
Granted there is one heck
of spike in the June of 1995 average, still I don't think the cyclopoids
numbers changed enough to say that their numbers have decreased in the
samples to any significant extant (this is where it would be useful to
know how to compute the margin of error in the data).
COPEPOD NAUPLII
No attempt was made to distinguish between cyclopoid
and calanoid copepods. I'm not sure if I could have told the difference
between the two even it the attempt had been made.
NAUPLII
NAUPLII (both cyclopoid and calanoid)
1993 1994 1995
1996 1997 1998 1999
2000
2001 2003
2004
Jan.
3.25 3.8 0.3
0.4
Feb.
.75 1.9 1.4
.29
Mar
9.5 1.4
.9 .25
Apr.
36.2 28
8.3 61.3 33.4
36.8 42.9
14.8 25.6
11.5
May
134 161.3 63.6
13.8 53.4 20.1
22.8
15.2 13.5
18.7
June 13
61.3 129.4 121.2
64.8 38.5 17.1
30.7
12.3 23.8
18.5
July
2.7 27.7 29.4
51 48.6
5.2 4.4
19
10
20.2 25.3
Aug. 4.5
49.6 70.4 30
15 9.8
14.4 6.7
5.9
Sept. 6.6
26 40.1
49.7 10 38.4
16.9 17.6
4.6
Oct.
4.8 10.2
13.1 44
8.8 14.8
7
10
0.4
Nov. 6.4
2.9 4.4
1.6 2.8
1.4 1.6
1.25
0
Dec
4.6 5.6
2.4 1.7 0.2
0.25 0.8
0
Yearly average 1994
1995 1996 1997 1998
1999 2000
2001
from Apr to Dec. 39.26 53.17
41.21 25.03 16.26 13.23
16.77 7.9
The graph below of the Nauplii is different
from those above in that I have included the month of April to the months
of May and June. The reason is that the Nauplii begin their population
increase a month earlier than the adults (which only makes sense). It would
appear that the number of nauplii found in the samples has gone down from
their high numbers in 1996 and before.

SUMMARY
Six or seven years seems
hardly sufficient to say there has been any change in the long term number
of copepods found in the Lake.. And having taken only somewhere between
500 and a 1000 samples seem paltry when one considers the sample size compared
to the size of the Lake, but its all I've got to work with. What makes
arriving at any kind of meaningful statement about changes in the number
of copepods foolhardy in my case is that I have no idea how these animals
actually disperse themselves in the water. I think I can safely say that
there is not always a random distribution of the animals, and not knowing
how often the animals swarm it becomes impossible to determine the value
or meaning of the samples taken.
NOTES ON THE YEAR 2001
Some data from 2001 for Cyclopoids
The cyclopoid copepods have for some reason
increased greatly in numbers in the month of May 2001. Why? I haven't a
clue. The number of these crustaceans is not the result of an increase
in one sample which is important in that a physical or biological may cause
aggregations of the copepods to appear in an area for a short time. Below
is a listing of the cyclopoids found in the samples for May and the beginning
of June 2001.
cyclopoids
May copepods
2
2
6
24
9
7
12
41
16
117
20
33
22
919
27
905
31
10
average 228.66 per 12.5 liter sample
June
3
116
6
1251
10
38
13
14